{"id":142567,"date":"2025-05-12T13:48:07","date_gmt":"2025-05-12T13:48:07","guid":{"rendered":"https:\/\/www.unipi.it\/ateneo\/organizzazione\/persone\/ugo-borello-129512-en\/"},"modified":"2026-04-01T04:33:51","modified_gmt":"2026-04-01T02:33:51","slug":"ugo-borello-129512-en","status":"publish","type":"unipi_persone","link":"https:\/\/www.unipi.it\/en\/about-us\/organisation\/people\/ugo-borello-129512-en\/","title":{"rendered":"Ugo Borello"},"content":{"rendered":"<p>Professore Associato &#8211; Dipartimento di Biologia | Tel.: 050 2211492 &#8211; Email: ugo.borello@unipi.it<\/p>\n","protected":false},"template":"","meta":{"unipi-persone-chiave":"129512","unipi-persone-nome":"Ugo","unipi-persone-cognome":"Borello","unipi-persone-email":"ugo.borello@unipi.it","unipi-persone-cv":"dal 2011 al 2018<br \/>\r\nRicercatore, Stem Cell and Brain Research Institute, INSERM U1208, Bron, France.<br \/>\r\n<br \/>\r\ndal 2008 al 2011<br \/>\r\nBorsista Post-Doc, CNRS, Institut Jacques Monod, Paris, France.<br \/>\r\n<br \/>\r\ndal 2002 al 2008<br \/>\r\nBorsista Post-Doc, University of California San Francisco, San Francisco USA.<br \/>\r\n<br \/>\r\ndal 2001 al 2001<br \/>\r\nProfessore a contratto, Corso di Istologia ed Embriologia Medica, Universit\u00e0 \"La Sapienza\" di Roma, II Facolt\u00e0 di Medicina e Chirurgia.<br \/>\r\n<br \/>\r\ndal 2001 al 2002<br \/>\r\nAssegnista di Ricerca; Universit\u00e0 \"La Sapienza\", Roma; Istituto di Istologia ed Embriologia Medica.<br \/>\r\n<br \/>\r\ndal 1997 al 2001<br \/>\r\nDottorato di Ricerca, Universit\u00e0 di Roma \"La Sapienza\", Istituto di Istologia ed Embriologia Medica.","unipi-persone-link-esterno-cv":"https:\/\/people.unipi.it\/ugoborelloit\/curriculum\/","unipi-persone-modalita-ricevimento":"","unipi-persone-luogo-ricevimento":"Unit\u00e0 di Biologia Cellulare e dello Sviluppo\nS.S. 12 Abetone e Brennero 4,\n 56127 Pisa\n\nemail: ugo.borello@unipi.it","unipi-persone-ore-ricevimento":"Tutti i giorni previo appuntamento via email.","unipi-persone-telefono":"050 2211492","unipi-persone-sitoweb":"","unipi-persone-cariche":"","unipi-persone-link-orario-lezioni":"https:\/\/unipi.prod.up.cineca.it\/calendarioPubblico\/matricola=038613","unipi-persone-link-registro-lezioni":"https:\/\/unimap.unipi.it\/registri\/registri.php?ri=038613&tmplt=principale.tpl","unipi-persone-settore-scientifico":"Anatomia, Biologia Cellulare e Biologia Dello Sviluppo Comparate BIOS-04\/A","unipi-persone-ruolo":"Professore Associato","unipi-persone-arpi":"https:\/\/arpi.unipi.it\/cris\/rp\/rp84826","unipi-persone-pubblicazioni":"<ul class=\"pubblicazioni-list\"><li><a href=\"https:\/\/arpi.unipi.it\/handle\/11568\/1347227\" target=\"_blank\" rel=\"noopener noreferrer\">Morpho-Functional Characterization and miRNA Profiling of the Retina in the 5xFAD Murine Model of Alzheimer&#039;s Disease<\/a> <span class=\"pub-details\">(Guidotti, Lorenzo; Lucchesi, Martina; Daghini, Eleonora; Amato, Rosario; Neri, Giuseppe; Corsi, Francesca; Marracci, Silvia; Gargini, Claudia; Borello, Ugo; Cammalleri, Maurizio; Dal Monte, Massimo; Casini, Giovanni - 2026 - 1.1 Articolo in rivista)<\/span><\/li><li><a href=\"https:\/\/arpi.unipi.it\/handle\/11568\/1343367\" target=\"_blank\" rel=\"noopener noreferrer\">Bidirectional interaction between protocadherin 8 and the transcription factor Dbx1 regulates cerebral cortex development<\/a> <span class=\"pub-details\">(Cwetsch, Andrzej W; Ferreira, Sofia; Delberghe, Elodie; Gilabert-Juan, Javier; Moreau, Matthieu X; Saillour, Yoann; Garc\u00eda-Bolufer, Pau; Calvo-Parra, Saray; Gonz\u00e1lez-Mart\u00ednez, Jose; Massoukou, Durcia; Borello, Ugo; Causeret, Fr\u00e9d\u00e9ric; Pierani, Alessandra - 2026 - 1.1 Articolo in rivista)<\/span><\/li><li><a href=\"https:\/\/arpi.unipi.it\/handle\/11568\/1304848\" target=\"_blank\" rel=\"noopener noreferrer\">Giuseppe Levi [1872-1965]<\/a> <span class=\"pub-details\">(Borello, Ugo; Crapanzano, Erica Lucia - 2025 - 1.1 Articolo in rivista)<\/span><\/li><li><a href=\"https:\/\/arpi.unipi.it\/handle\/11568\/1277727\" target=\"_blank\" rel=\"noopener noreferrer\">Comparative proteomic analysis of human vitreous in rhegmatogenous retinal detachment and diabetic retinopathy reveals a common pathway and potential therapeutic target<\/a> <span class=\"pub-details\">(Brighenti, Tommaso; Neri, Giuseppe; Mazzola, Marco; Tom\u00e9, Gabriele; Scalfati, Mariella; Peroni, Daniele; Belli, Romina; Zampedri, Elena; Tebaldi, Toma; Borello, Ugo; Romanelli, Federica; Casarosa, Simona - 2024 - 1.1 Articolo in rivista)<\/span><\/li><li><a href=\"https:\/\/arpi.unipi.it\/handle\/11568\/1271228\" target=\"_blank\" rel=\"noopener noreferrer\">RettDb: the Rett syndrome omics database to navigate the Rett syndrome genomic landscape<\/a> <span class=\"pub-details\">(Cillari, Nico; Neri, Giuseppe; Pisanti, Nadia; Milazzo, Paolo; Borello, Ugo - 2024 - 1.1 Articolo in rivista)<\/span><\/li><li><a href=\"https:\/\/arpi.unipi.it\/handle\/11568\/1271227\" target=\"_blank\" rel=\"noopener noreferrer\">Neural precursor cells rescue symptoms of Rett syndrome by activation of the Interferon ? pathway<\/a> <span class=\"pub-details\">(Frasca, Angelisa; Miramondi, Federica; Butti, Erica; Indrigo, Marzia; Balbontin Arenas, Maria; Postogna, Francesca M; Piffer, Arianna; Bedogni, Francesco; Pizzamiglio, Lara; Cambria, Clara; Borello, Ugo; Antonucci, Flavia; Martino, Gianvito; Landsberger, Nicoletta - 2024 - 1.1 Articolo in rivista)<\/span><\/li><li><a href=\"https:\/\/arpi.unipi.it\/handle\/11568\/1167887\" target=\"_blank\" rel=\"noopener noreferrer\">Axonal plasticity in response to active forces generated through magnetic nano-pulling<\/a> <span class=\"pub-details\">(Falconieri, Alessandro; De Vincentiis, Sara; Cappello, Valentina; Convertino, Domenica; Das, Ravi; Ghignoli, Samuele; Figoli, Sofia; Luin, Stefano; Catal\u00e0-Castro, Frederic; Marchetti, Laura; Borello, Ugo; Krieg, Michael; Raffa, Vittoria - 2023 - 1.1 Articolo in rivista)<\/span><\/li><li><a href=\"https:\/\/arpi.unipi.it\/handle\/11568\/1167905\" target=\"_blank\" rel=\"noopener noreferrer\">RETT SYNDROME OMICS DATABASE<\/a> <span class=\"pub-details\">(Borello, Ugo; Cill\u00e0ri, Nico; Milazzo, Paolo; Neri, Giuseppe; Pisanti, Nadia; Scalisi, Marta - 2022 - 5.10 Banca dati)<\/span><\/li><li><a href=\"https:\/\/arpi.unipi.it\/handle\/11568\/1151079\" target=\"_blank\" rel=\"noopener noreferrer\">D-aspartate oxidase gene duplication induces social recognition memory deficit in mice and intellectual disabilities in humans<\/a> <span class=\"pub-details\">(Lombardo, B.; Pagani, M.; De Rosa, A.; Nunziato, M.; Migliarini, S.; Garofalo, M.; Terrile, M.; D&#039;Argenio, V.; Galbusera, A.; Nuzzo, T.; Ranieri, A.; Vitale, A.; Leggiero, E.; Di Maio, A.; Barsotti, N.; Borello, U.; Napolitano, F.; Mandarino, A.; Carotenuto, M.; Heresco-Levy, U.; Pasqualetti, M.; Malatesta, P.; Gozzi, A.; Errico, F.; Salvatore, F.; Pastore, L.; Usiello, A. - 2022 - 1.1 Articolo in rivista)<\/span><\/li><li><a href=\"https:\/\/arpi.unipi.it\/handle\/11568\/1139411\" target=\"_blank\" rel=\"noopener noreferrer\">Centrin 2: a novel marker of mature and neoplastic human astrocytes<\/a> <span class=\"pub-details\">(Degl&#039;Innocenti, Elisa; Emanuele Poloni, Tino; Medici, Valentina; Recupero, Luca; Dell&#039;Amico, Claudia; Vannini, Eleonora; Borello, Ugo; Mazzanti, Chiara M.; Onorati, Marco; Teresa Dell&#039;Anno, Maria - 2022 - 1.1 Articolo in rivista)<\/span><\/li><li><a href=\"https:\/\/arpi.unipi.it\/handle\/11568\/1054702\" target=\"_blank\" rel=\"noopener noreferrer\">The Stemness Gene Mex3A Is a Key Regulator of Neuroblast Proliferation During Neurogenesis<\/a> <span class=\"pub-details\">(Naef, V.; De Sarlo, M.; Testa, G.; Corsinovi, D.; Azzarelli, R.; Borello, U.; Ori, M. - 2020 - 1.1 Articolo in rivista)<\/span><\/li><li><a href=\"https:\/\/arpi.unipi.it\/handle\/11568\/1026835\" target=\"_blank\" rel=\"noopener noreferrer\">Cortical Seizures in FoxG1 +\/- mice are accompanied by akt\/s6 overactivation, excitation\/inhibition imbalance and impaired synaptic transmission<\/a> <span class=\"pub-details\">(Testa, G.; Olimpico, F.; Pancrazi, L.; Borello, U.; Cattaneo, A.; Caleo, M.; Costa, M.; Mainardi, M. - 2019 - 1.1 Articolo in rivista)<\/span><\/li><li><a href=\"https:\/\/arpi.unipi.it\/handle\/11568\/933217\" target=\"_blank\" rel=\"noopener noreferrer\">SP8 transcriptional regulation of Cyclin D1 during mouse early corticogenesis<\/a> <span class=\"pub-details\">(Borello, Ugo; Berarducci, Barbara; Delahaye, Edwige; Price, David J.; Dehay, Colette - 2018 - 1.1 Articolo in rivista)<\/span><\/li><li><a href=\"https:\/\/arpi.unipi.it\/handle\/11568\/933214\" target=\"_blank\" rel=\"noopener noreferrer\">The logistics of afferent cortical specification in mice and men<\/a> <span class=\"pub-details\">(Borello, Ugo; Kennedy, Henry; Dehay, Colette - 2018 - 1.1 Articolo in rivista)<\/span><\/li><li><a href=\"https:\/\/arpi.unipi.it\/handle\/11568\/933010\" target=\"_blank\" rel=\"noopener noreferrer\">Radial derivatives of the mouse ventral pallium traced with Dbx1-LacZ reporters<\/a> <span class=\"pub-details\">(Puelles, L; Medina, L; Borello, U; Legaz, I; Teissier, A; Pierani, A; Rubenstein, Jlr - 2016 - 1.1 Articolo in rivista)<\/span><\/li><\/ul>","unipi-persone-didattica":"<ul class=\"registri-list\"><li>Citologia e istologia - corso b (cod. 071EE) per Laurea in Scienze biologiche<\/li><\/ul>","unipi-persone-codocenze":"<h4 class=\"mt-2\">Incarichi di co-docenza in moduli\/insegnamenti<\/h4><ul><li>Cognizione Molecolare della Cellula (Cod. 0006E) per Laurea Magistrale in Biologia molecolare e cellulare<\/li><li>Biologia Dello Sviluppo (Cod. 068EE) per Laurea in Scienze biologiche<\/li><\/ul>","unipi-persone-interessi-ricerca":"","_wpml_word_count":"13","_last_translation_edit_mode":"translation-editor","unipi-persone-dati-posizione":""},"organizzazioni":[3588],"class_list":["post-142567","unipi_persone","type-unipi_persone","status-publish","hentry","organizzazioni-dipartimento-di-biologia-409999-en"],"email":"ugo.borello@unipi.it","telefono":"050 2211492","content_plain":"Professore Associato &#8211; Dipartimento di Biologia | Tel.: 050 2211492 &#8211; Email: ugo.borello@unipi.it","photo_url":null,"ruoli_tags":[{"id":759,"name":"PA","slug":"pa"}],"yoast_head":"<!-- This site is optimized with the Yoast SEO plugin v27.0 - https:\/\/yoast.com\/product\/yoast-seo-wordpress\/ -->\n<title>Ugo Borello ~ UNIPI<\/title>\n<meta name=\"robots\" content=\"index, follow, max-snippet:-1, max-image-preview:large, max-video-preview:-1\" \/>\n<link rel=\"canonical\" href=\"https:\/\/www.unipi.it\/en\/about-us\/organisation\/people\/ugo-borello-129512-en\/\" \/>\n<meta property=\"og:locale\" content=\"en_US\" \/>\n<meta property=\"og:type\" content=\"article\" \/>\n<meta property=\"og:title\" content=\"Ugo Borello ~ UNIPI\" \/>\n<meta property=\"og:description\" content=\"Professore Associato &#8211; Dipartimento di Biologia | Tel.: 050 2211492 &#8211; Email: ugo.borello@unipi.it\" \/>\n<meta property=\"og:url\" content=\"https:\/\/www.unipi.it\/en\/about-us\/organisation\/people\/ugo-borello-129512-en\/\" \/>\n<meta property=\"og:site_name\" content=\"UNIPI\" \/>\n<meta property=\"article:publisher\" content=\"https:\/\/www.facebook.com\/unipisaofficial\/\" \/>\n<meta property=\"article:modified_time\" content=\"2026-04-01T02:33:51+00:00\" \/>\n<meta name=\"twitter:card\" content=\"summary_large_image\" \/>\n<script type=\"application\/ld+json\" class=\"yoast-schema-graph\">{\"@context\":\"https:\/\/schema.org\",\"@graph\":[{\"@type\":\"WebPage\",\"@id\":\"https:\/\/www.unipi.it\/en\/about-us\/organisation\/people\/ugo-borello-129512-en\/\",\"url\":\"https:\/\/www.unipi.it\/en\/about-us\/organisation\/people\/ugo-borello-129512-en\/\",\"name\":\"Ugo Borello ~ UNIPI\",\"isPartOf\":{\"@id\":\"https:\/\/www.unipi.it\/en\/#website\"},\"datePublished\":\"2025-05-12T13:48:07+00:00\",\"dateModified\":\"2026-04-01T02:33:51+00:00\",\"breadcrumb\":{\"@id\":\"https:\/\/www.unipi.it\/en\/about-us\/organisation\/people\/ugo-borello-129512-en\/#breadcrumb\"},\"inLanguage\":\"en-US\",\"potentialAction\":[{\"@type\":\"ReadAction\",\"target\":[\"https:\/\/www.unipi.it\/en\/about-us\/organisation\/people\/ugo-borello-129512-en\/\"]}]},{\"@type\":\"BreadcrumbList\",\"@id\":\"https:\/\/www.unipi.it\/en\/about-us\/organisation\/people\/ugo-borello-129512-en\/#breadcrumb\",\"itemListElement\":[{\"@type\":\"ListItem\",\"position\":1,\"name\":\"\",\"item\":\"https:\/\/www.unipi.it\/en\/\"},{\"@type\":\"ListItem\",\"position\":2,\"name\":\"Ugo Borello\"}]},{\"@type\":\"WebSite\",\"@id\":\"https:\/\/www.unipi.it\/en\/#website\",\"url\":\"https:\/\/www.unipi.it\/en\/\",\"name\":\"UNIPI\",\"description\":\"UNIPI\",\"publisher\":{\"@id\":\"https:\/\/www.unipi.it\/en\/#organization\"},\"potentialAction\":[{\"@type\":\"SearchAction\",\"target\":{\"@type\":\"EntryPoint\",\"urlTemplate\":\"https:\/\/www.unipi.it\/en\/?s={search_term_string}\"},\"query-input\":{\"@type\":\"PropertyValueSpecification\",\"valueRequired\":true,\"valueName\":\"search_term_string\"}}],\"inLanguage\":\"en-US\"},{\"@type\":\"Organization\",\"@id\":\"https:\/\/www.unipi.it\/en\/#organization\",\"name\":\"UNIPI\",\"url\":\"https:\/\/www.unipi.it\/en\/\",\"logo\":{\"@type\":\"ImageObject\",\"inLanguage\":\"en-US\",\"@id\":\"https:\/\/www.unipi.it\/en\/#\/schema\/logo\/image\/\",\"url\":\"https:\/\/www.unipi.it\/wp-content\/uploads\/Raggruppa-3020.svg\",\"contentUrl\":\"https:\/\/www.unipi.it\/wp-content\/uploads\/Raggruppa-3020.svg\",\"width\":176,\"height\":64,\"caption\":\"UNIPI\"},\"image\":{\"@id\":\"https:\/\/www.unipi.it\/en\/#\/schema\/logo\/image\/\"},\"sameAs\":[\"https:\/\/www.facebook.com\/unipisaofficial\/\",\"https:\/\/www.instagram.com\/unipisa\/\"]}]}<\/script>\n<!-- \/ Yoast SEO plugin. -->","yoast_head_json":{"title":"Ugo Borello ~ UNIPI","robots":{"index":"index","follow":"follow","max-snippet":"max-snippet:-1","max-image-preview":"max-image-preview:large","max-video-preview":"max-video-preview:-1"},"canonical":"https:\/\/www.unipi.it\/en\/about-us\/organisation\/people\/ugo-borello-129512-en\/","og_locale":"en_US","og_type":"article","og_title":"Ugo Borello ~ UNIPI","og_description":"Professore Associato &#8211; 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